| Hackelia | |
|---|---|
![]()  | |
| Hackelia diffusa | |
| Scientific classification  | |
| Kingdom: | Plantae | 
| Clade: | Tracheophytes | 
| Clade: | Angiosperms | 
| Clade: | Eudicots | 
| Clade: | Asterids | 
| Order: | Boraginales | 
| Family: | Boraginaceae | 
| Subfamily: | Boraginoideae | 
| Genus: | Hackelia Opiz (1839)  | 
| Type species | |
| Hackelia deflexa Opiz  | |
| Species | |
| 
 About 54 (see text)  | |
| Synonyms[1] | |
| 
 Austrocynoglossum Popov ex R.Mill. (1989)  | |
Hackelia (stickseeds) is a genus of plants in the borage family, Boraginaceae. It includes 54 species found in North America, western South America, temperate Eurasia, and Australia.[1] 12 species are native to California.[2]
The genus was named after Josef Hackel, a Czech botanist. The common name, stickseed, refers to the tendency of the barbed nutlets to stick to animal fur.[3]
Species
54 species are currently accepted:[1]
- Hackelia amethystina J.T.Howell - amethyst stickseed – California
 - Hackelia andicola (K.Krause) Brand – Peru
 - Hackelia bella (J.F.Macbr.) I.M.Johnst. - greater showy stickseed – northwestern California and southwestern Oregon
 - Hackelia besseyi (Rydb.) J.L. Gentry - Bessey's stickseed – Arizona, Colorado, New Mexico and Texas
 - Hackelia bhutanica R.R.Mill – eastern Nepal to northwestern Arunachal Pradesh
 - Hackelia brachytuba (Diels) I.M.Johnst. – Nepal to central China
 - Hackelia brevicula (Jeps.) J.L.Gentry - Poison Canyon stickseed – eastern California
 - Hackelia californica (A.Gray) I.M.Johnst. - California stickseed – northern California and west-central Oregon
 - Hackelia ciliata (Douglas ex Lehm.) I.M.Johnst. - British Columbia, Idaho, and Washington
 - Hackelia cinerea (Piper) I.M.Johnst. - gray stickseed – Washington, Idaho, and Montana
 - Hackelia cronquistii J.L.Gentry - Cronquist's stickseed – Oregon and Idaho
 - Hackelia cusickii (Piper) Brand - Cusick's stickseed – Oregon, northern California, and northwestern Nevada
 - Hackelia davisii Cronquist – Idaho
 - Hackelia deflexa (Wahlenb.) Opiz – northern Eurasia and northern North America
 - Hackelia difformis (Y.S.Lian & J.Q.Wang) Riedl – south-central China and Tibet
 - Hackelia diffusa (Douglas ex Lehm.) I.M.Johnst. - Spreading stickseed – British Columbia, Washington, Oregon
 - Hackelia floribunda (Lehm.) I.M.Johnst. - manyflower stickseed – western and central Canada and United States
 - Hackelia gracilenta I.M.Johnst. – Colorado
 - Hackelia heliocarpa (Brand) Brand – Mexico (Chihuahua)
 - Hackelia hendersonii (Piper) Brand – Washington, Oregon, and Montana
 - Hackelia hintoniorum (B.L.Turner) Sutorý – Mexico (Oaxaca)
 - Hackelia hirsuta (Wooton & Standl.) I.M.Johnst. – New Mexico
 - Hackelia hispida I.M.Johnst. - showy stickseed – Washington, Oregon, Idaho
 - Hackelia ibapensis L.M.Shultz & J.S.Shultz – Utah
 - Hackelia latifolia (R.Br.) Dimon & M.A.M.Renner – eastern Australia and Tasmania
 - Hackelia leonotis I.M.Johnst. – northeastern and central Mexico
 - Hackelia macrophylla (Brand) I.M.Johnst. – northern Pakistan and western Himalaya
 - Hackelia meeboldii Brand – western Himalaya
 - Hackelia mexicana (Schltdl. & Cham.) I.M.Johnst. – Mexico to Venezuela and Peru
 - Hackelia micrantha (Eastw.) J.L.Gentry - Jessica sticktight – western Canada and United States
 - Hackelia mundula (Jeps.) Ferris - pink stickseed – southwestern Oregon to central California
 - Hackelia murgabica Czukav. – Tajikistan
 - Hackelia nervosa (Kellogg) I.M.Johnst. - Sierra stickseed – California and northwestern Nevada
 - Hackelia obtusifolia R.R.Mill – Bhutan
 - Hackelia ophiobia R.L.Carr – southeastern Oregon, southwestern Idaho, and northern Nevada
 - Hackelia parviflora (K.Krause) Brand – Peru (Junín)
 - Hackelia patens (Nutt.) I.M.Johnst. - spotted stickseed – west-central United States
 - Hackelia pinetorum (Greene ex A.Gray) I.M.Johnst. – Arizona, New Mexico, Texas, and northern Mexico
 - Hackelia popovii Czukav. – Tajikistan
 - Hackelia rattanii (Brand) Brand – Mexico (Baja California)
 - Hackelia revoluta (Ruiz & Pav.) I.M.Johnst. – Colombia to northwestern Argentina
 - Hackelia sessilifructa (Y.S.Lian & J.Q.Wang) Ovczinnikova – Xinjiang
 - Hackelia setosa (Piper) I.M.Johnst. - bristly stickseed – northern California and southwestern Oregon
 - Hackelia sharsmithii I.M.Johnst. - Sharsmith's stickseed – central California and central Nevada
 - Hackelia skutchii I.M.Johnst. – Guatemala and Costa Rica
 - Hackelia stewartii I.M.Johnst. – northern Pakistan and western Himalaya
 - Hackelia stricta I.M.Johnst. – southwestern Mexico to Veracruz
 - Hackelia suaveolens (R.Br.) Dimon & M.A.M.Renner – eastern and southern Australia and Tasmania
 - Hackelia taylorii Harrod, Malmquist & R.L.Carr – Washington
 - Hackelia uncinata (Royle ex Benth.) C.E.C.Fisch. – northeastern Pakistan to Himalaya and southern China
 - Hackelia ursina (Greene ex A.Gray) I.M.Johnst. – southeastern Arizona, southwestern New Mexico, and northern Mexico (Sonora)
 - Hackelia velutina (Piper) I.M.Johnst. - velvet stickseed – north-central and central California and northwestern Nevada
 - Hackelia venusta (Piper) H.St.John - lesser showy stickseed – central Washington
 - Hackelia virginiana (L.) I.M.Johnst. - beggar's lice – central and eastern United States and eastern Canada
 
References
- 1 2 3 Hackelia Opiz. Plants of the World Online. Retrieved 17 June 2023.
 - ↑ Calflora Database: Hackelia species index in California. Calflora. Retrieved 17 June 2023.
 - ↑ Taylor, Ronald J. (1994) [1992]. Sagebrush Country: A Wildflower Sanctuary (rev. ed.). Missoula, MT: Mountain Press Pub. Co. p. 24. ISBN 0-87842-280-3. OCLC 25708726.
 
External links
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